The Polystichum Homepage

Welcome to the website for the hollyferns, members of the fern genus Polystichum and allies.
This website is maintained for the use of hollyfern enthusiasts and scholars
throughout the world by Dave Barrington at the University of Vermont in the United States.

Polystichum Homepage

Portrait of the Genus


Species List

Diversity Map

Interactive Keys


Cytology (PDF)

Barrington Web Page
(with publications)

Morphological Characters

Morphological Analysis

Lab Protocols

Current Working World Phylogeny for Polystichum
(128 species, four sequences)

This is the working Polystichum phylogeny currently used in the Barrington lab. To view the most recently published phylogeny (Driscoll & Barrington 2007), click here: Driscoll & Barrington 2007 Phylogeny


Please be in touch with me with your questions about and insights into the holly ferns -