209 schedule

CALENDAR AND ACTIVITIES FOR BIOLOGY OF FERNS, PLANT BIOLOGY 209
2023

Welcome to the 2023 Schedule for Biology of Ferns. This page and the powerpoints chnge all the time. I continue adapting the schedule to the new literature that I've encountered since I last offered the course and to new approaches to teaching the material.

The part that I have changed for this year is a darker shade of green; the part that remains to be changed is paler.

The powerpoints are all linked her, in capital letters and italic.

The readings are all on-line, linked below.

This is the version of March 20, 2023, at 5 pm

Sequence of topics
A. Morphology
B. Sterile Hybrids
C. Polyploids
D. Apomixis
E. Biogeography
F. Phylogeny This color is 8bb381

Calendar

DATE	TOPIC AND LINK TO PPTs	READINGS and REFERENCES (Link to full bibliography)
Jan. 17	Introduction, morphology 1: the fern life cycle, introduction and sporophyte MORPHOLOGY	Raven et. al. 2005, pp. 489-498
Jan. 19	sporophytes, meiosis, and spores
Jan. 22	gametophytes; vegetative and reproductive; Morphology more closely: sporophyte first part, the stem in part
Jan. 24	Morphology concluded; stem, leaf, and sorus
Jan. 26	Hybrids 1 - introduction to hybridization, intermediate morphology Hybrids 2 - using chromosome pairing to test hybrid hypotheses STERILE HYBRIDS	Barrington, 1986
Jan. 31	Hybrids 3 - using isozymes and DNA to test hybrid hypotheses, a case study with Polystichum concinnum x P. speciosissimum	Wagner et al. 1992 Rothfels et al., 2015. Rothfels et al. supplemental file Grimm et al. 2015 Wagner et al., 1978
*Feb. 2*	Hybrids 4 - examples from Dryopteris, the outer(and age-related) limits of hybrids
*Feb. 7*	Polyploidy 1 - Manton on Polystichum aculeatum POLYPLOIDS	Manton 1950, Ch. 6, pp. 88-92, Ch. 9, pp. 151-156
*Feb. 9*	Polyploidy 2 - Herb Wagner on Appalachian Asplenium, The Mechanics of Polyploidy	Wagner, 1954
*Feb. 14*	Polyploidy 3 - multiple origins	Sigel et al. 2014 Werth, 1985
*Feb. 16*	Polyploidy 4 - autopolyploidy spore size, flow cytometry, and genome size The origins of Polystichum braunii and Adiantum pedatum were not covered in lecture, but the slides for those segments are in the powerpoints.	Jorgensen and Barrington, 2017
*Feb. 21*	Apomixis - discovery of apomixis, apomictic Vermont ferns, Manton's scheme APOMIXIS	Manton, 1950 - chapter 10 on apogamy (apomixis)
*Feb. 23*	Origin of apomixis in our Vermont ferns: Pellaea The Braithwaite apomixis pathway Amanda Grusz's study of apomixis and polyploidy in three desert ferns	Grusz, 2009 Grusz, 2016 Braithwaite, 1964
Feb. 28	Cornopteris hybrids as possible indications of how apomixis originates. Apomictic species contributing sperm to new apomictic species Introduction to divergent evolution: divergence within species	Park et al. 2003
*Mar. 2*	Divergent evolution, continued: species stories—Haufler, Andrews, and Yatabe DIVERGENT EVOLUTION	Haufler et al., 2000 Yatabe et al., 2009
*Mar. 9*	Escape to the greenhouse
*Mar. 13-17*	Spring Break
Mar. 21	Biogeography 1: spore dispersal (ecological approach) BIOGEOGRAPHY 1	Barrington, 1993
. *Mar. 23*	Biogeography 2: spore dispersal (geographic approach) and Hawaii part 1
*Mar. 28*	Biogeography 3: Hawaii—, Taiwan, Continental Islands, Parameters of successful long-distance dispersal
Mar. 30	Biogeography 4: Biodiversity hot spots, endemic centers, and the origin of diversity BIOGEOGRAPHY 2
Apr. 4	Biogeography 5: vicariance and periglacial migration High polyploids on islands HIGH POLYPLOIDS
Apr. 6	Phylogeny 1 - the early groups, beginning with Osmundaceae Table of General Characters for the Ferns PHYLOGENY - THE EARLY GROUPS	PPG I 2016
Apr. 11	Phylogeny 2 - the early groups, continued
*Apr. 13*	Phylogeny 3 - the early groups, concluded (Cyatheales)
*Apr. 18*	Phylogeny 4 - the Polypodiales: early groups PHYLOGENY - THE RECENT GROUPS
*Apr. 20*	Phylogeny 5 - the recent groups, Eupolypods II	Schuettpelz & Pryer, 2009
Apr. 25	Phylogeny 6 - the recent groups, Eupolypods I
Apr. 27
May 2
May 4
May 12	Final Exam, 1:30 to 4:15