January 2004




David E. Capen

Wildlife and Fisheries Biology Program

Rubenstein School of Environment and Natural Resources

University of Vermont

Burlington, Vermont  05405

(802) 656-3007  david.capen@uvm.edu






University of Tennessee, B.S.F., 1969 (Forestry)

University of Maine, M.S., 1972 (Wildlife Management)

Utah State University, Ph.D., 1977 (Wildlife Science)




2002-Present: Research Professor, School of Natural Resources, University of Vermont

1999-2002: Professor, School of Natural Resources, University of Vermont

1999_2000: Visiting Scientist, National Parks and Wildlife Service, New South Wales, Australia (sabbatical leave)

1982-1999: Associate Professor, School of Natural Resources, University of Vermont

1993: Visiting Faculty Member, Department of Fisheries and Wildlife, Utah State University (sabbatical leave)

1987-1988: Acting Chair, Wildlife and Fisheries Biology Program, School of Natural Resources, University of Vermont

1986: Research Affiliate, Caesar Kleberg Wildlife Research Institute, Kingsville, Texas (sabbatical leave)

1976-1982: Assistant Professor of Wildlife Biology, School of Natural Resources, University of Vermont

1977-1980: Chair, Wildlife Biology Program, School of Natural Resources, University of Vermont

1973-1976: Research fellow; Welder Wildlife Foundation through Utah Cooperative Wildlife Research Unit

1972-1975: Teaching assistant; Department of Wildlife Science, Utah State University

1969-1971: Graduate research assistant; Maine Cooperative Wildlife Research Unit




1987-1989:  Editor, Wildlife Society Bulletin, an international, peer-reviewed journal published by The Wildlife Society.

1986-1987: Associate Editor, Wildlife Society Bulletin

1994-1999: Editorial Panel, Wildlife Society Bulletin

1995-1996: Editor, Northern goshawk and forest management in the southwestern United States.  Technical Review 96-2, The Wildlife Society. 19pp.




     The Wildlife Society                                   Wilson Ornithological Society

     Society of American Foresters                  Cooper Ornithological Society

     The Ecological Society of America          Waterbird Society

     American Ornithologists Union                 Society of Field Ornithology

     Society of Conservation Biology                            




Certified Wildlife Biologist, The Wildlife Society

Certified Forester, Society of American Foresters

Auditor, Forest Sustainability Certifications, FSC, SFI, ATF

Review for International Association of Fish and Wildlife Agencies: Proposal to Introduce Wild Turkeys into Nova Scotia

Professional Review Panel, USGS/BRD, Inventory and Monitoring

Numerous development-related wildlife consulting experiences in Vermont


FUNDED RESEARCH  (1994-2004)


Metapopulation Dynamics, decision-analysis modeling, and management of double-crested cormorants in the Lake Champlain Basin.  Lake Champlain Sea Grant.  2004-2005. (With T. Donovan and M. Richmond)


Do Conservation Easements Promote Sustainable Management of the Northern Forest?  North East State Foresters Association and Northeastern State Research Consortium, 2003-2004. (With P. Stokowski)


Ecological Reserves, Conservation Practices, and a Sustainable Working Landscape in New England’s Northern Forest.  Northeastern States Research Consortium, 2002-2005.  (With C. Danks)


Watershed-based Designs for Forest Conservation in an Ecologically Diverse Region of Northern New England.  McIntire-Stennis Cooperative Forest Research Funds, 2002-2004.


A Design for Conservation of Biological Diversity Based on Physical Diversity of the Landscape. McIntire-Stennis Cooperative Forest Research Funds. 1997-2001.


Comprehensive Conservation Planning for Lake Umbagog National Wildlife Refuge.  The Trust for Public Lands, 2002-2004.


Comprehensive Conservation Planning Missisquoi National Wildlife Refuge.  The Trust for Public Lands, 2002-2003.


Assessing the Relationship Between Landscape Diversity and Biological Diversity in an Upland Forested Ecosystem. Vermont Monitoring Cooperative. 2001-2002.


Diet, Movement, and Dispersal patterns of Double-crested Cormorants in Lake Champlain. Lake Champlain Sea Grant, U.S. Fish and Wildlife Service, Vermont Fish and Wildlife Department. 2001-2003. (With T. Donovan and D. Parrish)


The Lewis Creek Watershed: Demonstration of GIS Applications for Natural Resource Assessment, Development Analysis, and Conservation Education in Local Communities. USDA Special Research Grant. 1999-2000.


Spatial Models for Land Conservation Planning: Vermont Biodiversity Project.  Orton Family Foundation, The Nature Conservancy, Vermont Land Trust, others. 1997-2003.


A Habitat-based Analysis of Avian Diversity on the Ethan Allen Firing Range.  Vermont Military. 1998-1999.


Foraging Behavior and Resource Use by Neotropical Migratory Birds in a Shade Coffee Plantation in Belize.  University of Vermont, UCRS, SUGR & FAME, International Advisory Council; Wildlife Conservation Society


A Spatial Data Base to Support Planning for Conservation Lands in the Connecticut River Valley of Vermont. Silvio Conte National Fish and Wildlife Refuge Challenge Cost Share Program. 1997.


Landscape-level Prediction of Black-backed Woodpecker Habitat in Northeastern Vermont.  Silvio Conte National Fish and Wildlife Refuge Challenge Cost Share. 1997.


Planning for Conservation Lands: Southern Vermont Pilot Project. Vermont Department of Fish and Wildlife



White Mountain National Forest Wildlife Monitoring Program: Analyses of Data from Bird Surveys on Permanent Plots.  White Mountain National Forest. 1995-1997,


Population Dynamics and Food Habits of Double-crested Cormorants on Lake Champlain. U.S. Fish and Wildlife Service. 1996.


A Spatially Explicit Classification System for Atlantic Salmon Habitat on National Forests in New England.  Northeastern Forest Experiment Station, USDA Forest; Green Mountain National Forest. 1996-1998.


Development and Analysis of an Ecological Database for Multiple-use Planning on the Green Mountain National Forest.  Green Mountain National Forest and National Biological Service. 1992-1998.


Development of a Classification System for Wetlands on the Green Mountain National Forest.  Green Mountain National. 1995-1996.


Gap Analysis of Biological Diversity in Vermont and New Hampshire.  National Biological Service, USGS/BRD. 1993-1999.


Landscape-level Modeling of  Habitat Heterogeneity and Vertebrate Diversity. McIntire-Stennis Cooperative Forest Research Funds. 1995-1997.


Spatially-dependent Models of Avian Habitat Diversity on the Green Mountain National Forest.  Northeastern Forest Experiment Station. 1994-1996.


Biological Diversity in Vermont, New Hampshire, and the Northeastern U.S.: An Evaluation of Potential Species Richness in Relation to Habitat and Existing Reserves. U.S. Fish and Wildlife Service, National Biological Service. 1992-1994. (With K. Williams)



University of Vermont--1992-2003


WFB 388: Reserve Design (1 credit, 4-10 graduate students), Spring 2001, 2003

NR 224: Conservation Biology (3 credits 20-32 students), Spring 1995, 1996, 1997,1998, 1999, 2001, 2002 (Team-taught, 50%)

WFB 286: Special Topics--Landscape Metrics (1 credit, 12 graduate students), Spring 1995

WFB 286: Special Topics--Wildlife Habitat Analysis (1-3 credits, 9 graduate students), Fall 1993

WFB 174: Principles and Practices of Wildlife Management (3 credits, 80-140 students), Fall 1992-1994 (SNR Core); (40-60 students) Fall 1995, 1996, 1997, 1998, 1999

WFB 130: Ornithology (3 credits, 50-70 students) Spring 1992, 1995, 1996, 1997, 1998, 1999

WFB 131: Field Ornithology (2 credits, 24 students), Summer 1992, 1994, 1998

WFB 150: Wildlife Habitat and Population Measurements (2 credits, 20 students), Summer 1992,  (Team taught: 25%), (1 credit, 12 students), Summer 1998, 1999, 2000, 2001, 2002, 2003 (Team taught: 50%)


SERVICE (1994-2004)


Research and Scholarship


Member, Committee of Scientists, White Mountain National Forest, 1989-1997; Chair, Subcommittee on Birds

Peer Reviewer, Draft Spotted Owl Recovery Plan (800+ pages), U.S. Fish and Wildlife               Service/The Wildlife Society

Member, Research Working Group, Neotropical Bird Conservation Program, 1991-1995;  Chair, Technical Review Subcommittee

Vice-chair, Vermont Neotropical Migratory Bird Working Group, 1993-1994

Referee and Reviewer:  Journal of Wildlife Management, Wildlife Society Bulletin, Wildlife Monographs, Northeast Wildlife, Auk, Condor, Ecology, Conservation Biology, Landscape Ecology, National Science Foundation, Forest Ecology and Management, Canadian Journal of Forest Research, Western Journal of Applied Forestry, Wetlands, Colonial Waterbirds, Geographical and Environmental Modelling, USDA Forest Service--about 20 journal manuscripts  per year.




Member, Development Committee, Endowment Fund, The Wildlife Society, 2002-2003

Member, Honorary Membership and Awards Committee, The Wildlife Society, 2002-2004

Member, Editor Selection Committee, The Wildlife Society, 1996

Chair, Editor Selection Committee, Northeast Section, The Wildlife Society, 1994-1995

Member, Publications Awards Committee, The Wildlife Society, 1993-1994

Member, Program Committee, Northeast Fish and Wildlife Conference, 1994-1995

Member, Publications Committee, The Wildlife Society, 1994-1995




Member, Graduate Standards Committee, School of Natural Resources, 2003-2005

Member and Chair, Faculty Standards Committee, School of Natural Resources, 2000-2003

Chair, Faculty  Search Committee,  School of Natural Resources, 2000-2001

Member, Nominations Committee, UVM Senate, 1997-1999

Member, Executive Committee, Biological Sciences Curriculum, College of Agriculture and Life Sciences, UVM

Chair, Faculty Standards Committee, School of Natural Resources, 1995-1996

Co-chair, Faculty Search Committee, School of Natural Resources, 1995

Member, Faculty Affairs Committee, UVM Senate 1991-1995


Community and State


Member, Board of Directors, Vermont Center for Geographic Information, 2000-2004 (appointed by Governor)

Co-chair, Vermont Ecomapping Roundtable, 1997-1998

Member, Steering Committee, Scientific Advisory Committee, Technical Working Group, Vermont Biodiversity Project, 1996-2003; Chair, 2003-2004

Chair, Geological and Landscape Diversity Subcommitee, and Member, Scientific Advisory Group, New Hampshire Ecological Reserve Selection Committee, 1995-1998

Member, Scientific Advisory Group for Birds, Vermont Endangered Species Committee, 1988-2004

     Member, Conservation Commission, Town of Huntington, 1991-1994




Coordinator, Techniques for Landscape Analysis.  Two-day continuing education workshop for fish and wildlife biologists from state and federal agencies, March 1997.

Coordinator, Desktop GIS and Conservation Land Planning. One-day workshops for conservation lands planners, January 1997; for state foresters, April 1997.

Coordinator, GIS and Remote Sensing Applications for Wildlife and Fisheries; continuing education for mid-level professionals in U.S. Forest Service, March 1995. 


BIBLIOGRAPHY (1994-2004)


Refereed Publications


Hughes, M.E., and D.E. Capen.  Effects of local- and landscape-scale variation on the distribution of three area-sensitive forest songbirds in Vermont.  Condor.  In review.


Wells, D.B., and D.E. Capen.  A comparison of two methods for assessing songbird breeding productivity.  Journal of Field Ornithology. In review.


Wells, D.B., and D.E. Capen.  Powerline rights-of-way as breeding habitat for three shrubland bird species in southeastern New Hampshire: source, sink, or stronghold? Wildlife Society Bulletin.  In review.


Carlson, B.D., D. Wang, D.E. Capen, and E. H. Thompson.  2003. Landscape diversity units as predictors of forest community distributiona tool for regional conservation planning. J. Nature Conservation. In press.


Hurley, J.M., C. Ginger, and D.E. Capen. 2002. Property concepts, ecological thought, and ecosystem management.  Society and Natural Resources 15:297-315.


MacFaden, S.W., and D.E. Capen.  2002. Avian habitat relationships at multiple scales in a New England forest. Forest Science: 48(2): 243-253.


Ortega, Y.K., and D.E. Capen. 2002. Roads as edges: effects on birds in forested landscapes.  Forest Science 48(2): 381-390.


Coker, D.R., and D.E. Capen. 2000. Distribution and habitat associations of brown-headed cowbirds in the Green Mountains of Vermont.  Pages 236-243 in J.N.M. Smith, T.L. Cook, S.I. Rothstein, ,S.K. Robinson, and S.G. Sealy , Editors. Ecology and management of cowbirds and their hosts. University of Texas Press, Austin.


Capen, D.E., C. E. Ferree, and E. W. Buford. 1999. Landscape diversity as the basis for a reserve design initiative in Vermont.  Gap Analysis Bulletin 8:48-50.


Ortega, Y. K., and D. E. Capen. 1999.  Effects of forest roads on ovenbird habitat quality in a forested landscape.  Auk 116: 937-946.


Fowle, M. R., D. E. Capen, and N. J. Buckley. 1999. Population growth of double-crested cormorants in Lake Champlain.  Northeast Wildlife 54: 25-34.


Buford, E. W., and D. E. Capen.  1999. Abundance and productivity of forest songbirds in a managed, unfragmented forest.  Journal of Wildlife Management 63(1): In press.


Capen, D.E. 1999. Science Review: Northern Forest Lands Assessments. Pages 214-217 in K.N. Johnson, F.J. Swanson, M. Herring, and S. Greene, Editors. Bioregional assessments: science at the crossroads of management and policy. Island Press, Wash., D.C.


Germaine, S. S., S. H. Vessey, and D. E. Capen. 1997.  Effects of small forest openings on the breeding bird community in a Vermont hardwood forest.   Condor 99(3): 708-718.


Rykken, J. J., D. E. Capen, and S. Mahabir.  1997. Ground beetles (Coleoptera: Carabidae) as indicators of land type diversity in the Green Mountains of Vermont.  Conservation Biology 13(2): 522-530.


Buford, E. W., D. E. Capen, and B. K. Williams.  1996.  Distance sampling to estimate fledgling density of forest birds. Canadian Field-Naturalist 110(4): 1-7.


Coker, D. R., and D. E. Capen.  1995. Landscape-level habitat use by  brown-headed cowbirds in Vermont.  Journal of Wildlife Management 59(4): 631-637.


Lent, R. A., and D. E. Capen.  1995.  Effects of small-scale habitat disturbance on the ecology of  breeding birds in a Vermont (USA) hardwood forest.  Ecography 18(2): 97-108.


Other Publications


Richards, Z., and D.E. Capen. 2001. An assessment of management indicator species and habitat distribution models for monitoring viability of vertebrates on the Green Mountain National Forest.  Unpublished report submitted to Green Mountain National Forest. 29p.


Capen, D.E., E.W. Buford, and D. G. Williams (Contributing authors). 1998. An assessment of biodiversity of New Hampshire with recommendations for conservation actions.  M. Stevens, editor.  New Hampshire Chapter, The Nature Conservancy, Concord.


McFaden, S. W., D. E. Capen, and S. Fay.  1995.  An analysis of spatially-referenced data to identify potential research natural areas on the White Mountain National Forest.  Report submitted to White Mountain National Forest. 14 pp. 


Selected Presentations, Posters, Seminars


Capen, D.E.  Trials and tribulations of conservation planning in Vermont.  Invited Seminar, University of Maine, Department of Wildlife Ecology, 17 November 2003. Orono ME.


Capen, D.E. Landscape-level management of forests in New England: Why.  Invited paper, New England Society of American Foresters, 18 March 2003, Burlington, Vt.


Capen, D.E. The Vermont Biodiversity Project: conservation design based on landscape diversity.  Invited paper. International Association for Landscape Ecology--U.S. Regional Association. 25 April 2002, Lincoln, Nebraska.


Capen, D.E. Results of GAP analysis for Vermont and New Hampshire. Invited plenary address. Thinking big: landscape-level conservation in New Hampshire. 9 January 2002. Concord NH.


Capen, D.E. Systematic conservation planning to prevent fragmentation of habitat and populations.  Invited plenary address, Portuguese Ecological Society, 3 November 2001, Lisbon, Portugal.


Capen, D.E. Landscape diversity as a surrogate for biological diversity on national forests in New England.  Invited presentation. The Wilderness Society’s National Forest Activist Training. 23 February 2001, Fairlee VT.


Capen, D.E. Effectiveness of landscape surrogates for predicting biological diversity. Invited paper. Annual Conference of The Wildlife Society, 12-16 September 2001, Nashville TN.


Capen, D.E. Landscape diversity as the basis for conservation planning: examples from Vermont.  Invited presentation.  New England Society of American Foresters. 14 March 2001, Portland, ME.


Capen, D. E. Selection and design of ecological reserves: the Vermont Biodiversity Project.  Invited paper, Biodiversity in Maine: Issues and Opportunities.  Orono, Me, 20 November 1998.


Capen, D. E. Edges, openings, and bird habitat on the Green Mountain National Forest.  Invited seminar, University of Maine, Department of Wildlife Ecology, 26 October 1998.


Capen, D. E. Ecological reserve design in New England:  a comparison of science-based programs in three states.  Contributed poster, Annual Meeting, Society of Conservation Biology, Sydney, Australia, 13-16 July 1998.


Capen, D. E., T. L. Onega, E. W. Buford, D. G. Williams, and S. R. Magill.  Spatial modeling of black bear habitat in Vermont and New Hampshire.  Contributed poster, 11th International Conference on Bear Research and Management.  Gatlinburg, Tenn. 19-24 April 1998.


Capen, D. E. Growth of double-crested cormorant populations in Lake Champlain.  Invited presentation at workshop: Cormorant management in the Northeast.  Glenns Falls, N.Y. 13-15 January 1998.


Capen, D. E., E. W. Buford, and D. G. Williams.  Ecological reserve design in New England: data-rich gap analysis.  Contributed paper: 7th Annual Gap Analysis Principal Investigator's Meeting, Reston, Va. 4-8 August, 1997.


Capen, D. E.  (1) Gap analysis in New England, and (2) Discussion of  need for a regional assessment of biodiversity and regional design for ecological reserves. Invited presentations. Science-based efforts to design ecological reserves in northern New England.  Pinkham Notch, N.H., 24 September 1996.


Capen, D. E. Forest fragmentation and its impacts on habitat conditions for landbirds: a bi-state and regional perspective. Invited presentation.  New Hampshire/Vermont Bird Conservation Conference, Fairlee Vt, 12 April 1997.


Bove, J. R., and D. E. Capen.  Ecological land classification on the Green Mountain National Forest. Contributed poster. New England  Society of American Foresters, Portland, Me. 12-14 March 1997.  (Named "Second Best" Poster)


Capen, D. E.  Is there need for a regional assessment of biodiversity and a regional design for ecological reserves?  Invited presentation.  Science-based efforts to design ecological reserves in northern New England.  Pinkham Notch, N.H. 24 September 1996.


Capen, D. E.   Gap analysis in Vermont and New Hampshire and the appropriate scale for biodiversity monitoring.  Invited presentation. Landscape-level studies in New England: where are we and where are we going?  Appalachian Mountain Club Conference, Gorham, N. H., 28-29 September 1995.


Capen, D. E.  Songbirds in the Green Mountain National Forest: when does forest disturbance become forest fragmentation?  Invited paper. Songs of the deep woods. Conference sponsored by National Audubon Society. Stowe, Vt., 27 April 1995.


Capen. D. E.  The landscape-level approach to wildlife habitat studies.  Invited seminar. Department of Natural Resources, University of New Hampshire, 21 February 1995.


Capen, D. E.  Landscape-level habitat assessment: more statistics, more software, more pitfalls. Invited paper. Annual Conference, The Wildlife Society,  Albuquerque, N.M., 25 September 1994.


Capen, D. E. The peer-review process: assuring good science in print.  Invited paper. Northeast Fish and Wildlife Conference. Burlington, Vt., 3 May 1994.