The Polystichum Homepage

Welcome to the website for the hollyferns, members of the fern genus Polystichum and allies.
This website is maintained for the use of hollyfern enthusiasts and scholars
throughout the world by Dave Barrington at the University of Vermont in the United States.

Polystichum Homepage

Portrait of the Genus

Taxonomy

Species List

Diversity Map

Interactive Keys

Phylogeny

Cytology (PDF)

Barrington Web Page
(with publications)

Morphological Characters

Morphological Analysis

Lab Protocols



Key to Regions

01 Circumboreal
02 Eastern Asiatic
03 North American Atlantic
04 Rocky Mountain
05 Atlantic Islands
06 Mediterranean
07 Saharo-Arabian
08 Irano-Turanian
09 Madrean
10 Guineo-Congolian
11 Uzabara-Zululand
12 Sudano-Zabezian
13 Karoo-Namib
14 St Helena and Ascension
15 Madagascar & Mascarenes
16 Indian
17 Indochinese
18 Malesian
19 Fijian
20 Polynesian
21 Hawaiian
22 Neocaledonian
23M Caribbean - Mexico
23W Caribbean - West Indies
24 Guayana Highlands
25 Amazonian
26 Brazilian
27 Andean
28 Cape
29 Northeast Australia
30 Southwest Australian
31 Central Australian
32 Fernándezian
33 Chile-Pategonian
34 Subantarctic Islands
35 Neozeylandic

Current Names in Polystichum

This is the most inclusive list of possible species in the genus Polystichum; there are currently 343 names on the list, all of which have been recognized in at least one floristic or systematic work. In some cases, names have been excluded when recognized workers on the genus have made cogent arguments for their synonymy with other species. The numbers in brackets are literature references (a literature references page is forthcoming; ask for the literature citation in the meantime).

Regions are indicated by a numerical code; the key to this code is on the left side of the web page, underneath the navigation links. Regions are those of Takhtajan, 1986. Floristic Regions of the World. University of California Press, Berkeley CA. Some regions are subdivided further. Region 23 is divided into 23M (mainland Central America including tropical Mexico) and 23W (West Indies, mostly Greater Antilles).

Quality of species concept and thus number of species differs with region. In some regions (e.g. 11, 12, 29, 35), there have been modern systematic treatments based on non-morphological inquiries that have improved species concepts, so the quality is high. In others (e.g. 2, 18, 25), there has been no work on species concepts using modern tools, so species concepts are very much alpha-taxonomic. In the latter regions, the species number is almost certainly too high.

Improvements to this list are welcome.

Current Name Basionym Region
Polystichum acanthophyllum (Franch.) H. Christ [49] [57] Aspidium acanthophyllum Franch., Bull. Soc. Fr. 32.281885 02 16
Polystichum acrostichoides(Michx.)Schott, Gen. Fil. 1834 Nephrodium acrostichoides Michx., Fl.Bor.Am.2:267. Paris.Levrault.1803 03 09
Polystichum aculeatum (L.) Schott Polypodium aculeatum L., Sp. Plantarum 2:1090.1753. 01 02 06
Polystichum acutidens H. Christ Polystichum acutidens H. Christ, Bulletin de l'Académie internationale de géographie botanique 11(153-154): 259-260.1902 02 16 17 18
Polystichum acutipinnulum Ching & K.H. Shing [27] [295] Polystichum acutipinnulum Ching & K.H. Shing, Wuyi Sci. J. 1:91981. 02
Polystichum adungense Ching & Fraser-Jenk. ex H.S.Kung & L.B.Zhang [25] [295] Polystichum adungense Ching & Fraser-Jenk. ex H.S.Kung & L.B.Zhang, Acta Phytotax. Sin. 32(4):370. Fig. 1.1-1.61994. 02
Polystichum alaskense Maxon [db] Polystichum alaskense Maxon, Amer. Fern Journ. 8:35.1918. 04
Polystichum alatawshanicum C.Y.Yang [25] Polystichum alatawshanicum C.Y.Yang, Fl. Xinjiangensis 1:302.1992 02
Polystichum albomarginatum Kessler& A.R.Sm. [321] Polystichum albomarginatum M. Kessler & A.R.Sm., Brittonia 57(3): 218. f. 5A.2005. 27
Polystichum alcicorne (Baker) Diels [0] [166] Polypodium alcicorne Baker, JoB. 1888. 2291888 02
Polystichum alfaroi(H. Christ)Barrington [257, 258] Polystichum aculeatum (L.) Roth var. alfarii H. Christ, 23M
Polystichum alpinum Rosenst. [2] Polystichum alpinum Rosenst., Fedde Repert. 12:171.1913. 18
Polystichum altum Ching ex L.B. Zhang & H.S. Kung [166] [295] Polystichum altum Ching ex L.B. Zhang & H.S. Kung, Acta Phytotaxonomica Sinica 36(5):465ff 1998 & 37(1):82ff. 19991999. 02
Polystichum ambiguum Maxon [1] Polystichum ambiguum Maxon, Contr. USNat.Herb. 16:49 t.27.1912. 23W
Polystichum amboroense Kessler & A.R.Sm. [321] Polystichum amboroense M. Kessler & A.R.Sm., Brittonia 57(3): 224. f. 5D.2005. 27
Polystichum ammifolium (Poir.) C.Chr. Polypodium ammifolium Poir., Enc. 5.5541804 15
Polystichum andersonii Hopkins [2] Polystichum andersoni Hopkins, Amer. Fern Journ. 3:116 t. 9 1913. 04
Polystichum andinum Phil. [175] Polystichum andinum Phil., Linn. 29. 1081857 33 34
Polystichum anomalum (Hk. & Arn.) J.Sm. [57] Polypodium anomalum Hook. & Arn., JoB. 8. 360. t. 11.1856. 02 16 17
Polystichum archboldii Copel. [4] Polystichum archboldii Copel., Univ. Calif. Publ. Bot. 18:221.1942. 18
Polystichum assurgentipinnum W.M. Chu & B.Y. Zhang [166] Polystichum assurgentipinnum W.M. Chu & B.Y. Zhang, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):224. t. 41 f. 1-3 (p. 135). 2001. 02
Polystichum atkinsonii Bedd. [0] [10] [21 [57] 166] [295] Polystichum atkinsoni Bedd., Ferns br. Ind. suppl. 14 t. 362.1876. 02 16
Polystichum attenuatum Tagawa and Iwatsuki [db] Polystichum attenuatum Tagawa & Iwatsuki, Acta Phytotax. Geobot. 23:113, f.9.1968. 02 16
Polystichum auriculum Ching [4] [166] Polystichum auriculum Ching, Bull. Fan. Mem. Inst Biol. ser.2. 1:309.1949. 02
Polystichum auritum(Fée)Yatskiveych [27] Phanerophlebia aurita Fée, Cr.Vasc.Bres. 2:70, t.100.f.1.1872-3 26
Polystichum australiense Tindale [4] Polystichum australiense Tindale, S.Aust. Nat. 28: 31. f. 1. 1954. 29
Polystichum australium Copel. [db] Polystichum australium Copel., Occ. Pap. Bishop Mus. 14:57. t.8.1938. 20
Polystichum bachii M. Kessler& A.R.Sm. [321] Polystichum bachii M. Kessler & A.R.Sm., Brittonia 57(3): 214. f. 2A.2005. 27
Polystichum bamlerianum Ros.[1] Polystichum bamlerianum Ros., Fedd Repert. 10:330.1912. 18
Polystichum baoxingense Ching & H.S. Kung [27] [166] Polystichum baoxingense Ching & H.S. Kung, Acta Bot. Boreal-Occid. Sin. 9(4):271.1989. 02
Polystichum biaristatum (Blume) T. Moore [9] [49] [57] [117] Aspidium biaristatum Blume, Enum. 164.1828. 02 16 17 18
Polystichum biegemmatum Ching ex L.L. Xiang Polystichum bigemmatum Ching ex L.L. Xiang, Chenia 304: 133. f. 1-4. 1997. 02
Polystichum bifidum Ching [4] Polystichum bifidum Ching, Icon. Fil. Sin. 5: t. 242.1958. 02
Polystichum bissectum C.Chr. [1] Polystichum bissectum C.Chr., Ind. Suppl. 64.1913. 02
Polystichum bolanicum Rosenst. [2] Polystichum bolanicum Rosenst., Fedde Repert. (Repert. Spec. Nov. Regni Veg.)12:170.1913. 18
Polystichum bonseyi W.H.Wagner & R.W.Hobdy [25] Polystichum bonseyi W.H.Wagner & R.W.Hobdy, Contr. Univ. Michigan Herb. 19:75.1993. 21
Polystichum brassii Copel. [4] Polystichum brassii Copel., Philip. Journ. Sc. 78:394. t.2. 1951. 18
Polystichum braunii (Spenner) Fée [47] Aspidium braunii Spenner, Fl. Friburg 1,9 t. 21825. 02
Polystichum bulbiferum Barrington [27] Polystichum bulbiferum Barrington, Ann. Missouri Bot. Gard. 76(2):371.1989. 27
Polystichum calderonense Proctor [27] Polystichum calderonense Proctor, Mem. New York Bot. Gard. 53:273.1989. 23W
Polystichum californicum (Eaton) Diels Aspidium californicum Eaton, Pr. Amer. Ac. 6. 555. 1865. 09
Polystichum capillipes (Bak.)Diels [0] [57] [166] Aspidium capillipes Baker, JoB. 1888:228.1888 02
Polystichum caudatum (Sw.)Fée ? Aspidium freyreissii Wikstrom, "Nya eller mindre Kandz arter of Ormbunkar (Filices)."1826 26
Polystichum caudatum (Sw.)Fée [0, raised to species] Aspidium caudatum Sw., Vet.Akad.Handl.1817:62.[=Svenska Vet. Akademie Handingar]1817. 26
Polystichum caudescens Dutra Polystichum caudescens Dutra, An.Prim. Reun. Sul-Amer. Bot. 2:49.(Fl.Pter. do Rio Gr.do S.1940 26
Polystichum chaparensis Kessler& A.R.Sm. [321] Polystichum chaparensis M. Kessler & A.R.Sm., Brittonia 57(3): 225. f. 7.2005. 27
Polystichum cheilanthoides Copel. [4] Polystichum cheilanthoides Copel., Univ. Calif. Publ. Bot. 18:221. 1942. 18
Polystichum chilense(Christ)Diels [0 raised to species] Polystichum aculeatum (L.)Roth var. chilense H.Christ, Ber. Schw. Bot. Ges. 3:39.1893. 33
Polystichum chingae Ching [4] Polystichum chingae Ching, Icon. Fil. Sin. 4:t.182.1937. 02
Polystichum christianae (Jenm.)Und. & Maxon in Maxon Aspidium christianae G.S.Jenman, Bull.Bot.Dept.Jam. n.s.2:285.1895. 23W
Polystichum christii Ching [3] Polystichum christii Ching, Bull. Fan Mem. Inst. 2:192 t. 7. 1931. 02
Polystichum chunii Ching [3] [57] Polystichum chunii Ching, Sinensia 1:2., Ic. Fil. Sin. t. 13.1929. 02
Polystichum cochleatum (Klotzsch)Hieron. Polypodium cochleatum Klotzsch, Linn.20:388.1847 27 PAL
Polystichum concinnum Lellinger ex Barrington [27] Polystichum concinnum Lellinger ex Barrington, Ann. Missouri Bot. Gard. 76(2):367. f.2.1989. 23M
Polystichum congestum M. Kessler& A.R.Sm. [321] Polystichum congestum M. Kessler & A.R.Sm., Brittonia 57(3): 212. f. 2B.2005. 27
Polystichum controversum(Hance)Ching [3] Aspidium controversum Hance, Annales de Sci. naturelles: botanique IV.18:235.1862. 02
Polystichum copelandii Christ [1] Polystichum copelandii H. Christ, Phil. Journ. Sci.Bot. 2:157.1907. 18
Polystichum costularisorum Ching ex W. M. Chu and Z. R. He Polystichum costularisorum Ching ex W. M. Chu and Z. R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):224.2001. 02
Polystichum craspedosorum (Maxim.)Diels [0] [21] [34] [12] [47] Aspidium craspedosorum Maxim., Bull. Ac. St. Pétersb. 15:231. Mél. Biol. 7:341.[21]1871. 02
Polystichum crassinervium Ching ex W. M. Chu & Z. R. He [166] Polystichum crassinervium Ching ex W. M. Chu & Z. R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):224. t. 35 f. 1-4 (p. 122).2001. 02
Polystichum crinigerum(C.Chr.)Ching [4] [166] [295] Dryopteris crinigera C.Chr., Ind. Fil. Suppl. 1:31.1913. 02
Polystichum crinulosum(Desv.)J.P.Roux [25] Aspidium crinulosum Desvaux, Prod.249.1827. 15
Polystichum cuneatiforme Ching ex W.M. Chu & Z.R. He [166] Polystichum cuneatiforme W.M. Chu & Z. R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):1772001 02
Polystichum cuneatum Nakaike[27] Polystichum cuneatum Nakaike, Bull. Nat. Sci. Mus. Tokyo, B 4(3):128.1978. 18
Polystichum cyclolobum C.Chr. in Lév.[2] [57] Polystichum cyclolobum C.Chr. in H.Lév., Cat. Pl. Yun-Nan 111.1916. 02
Polystichum cystostegia (W.J. Hooker) J.B. Armstrong [0] [176] [218] Aspidium cystostegium W.J.Hooker, Sp. Fil.. 4:26. t.227.1862 35
Polystichum daymanense Nakaike [5] Polystichum daymanense Nakaike, Bot. Exped. Papua New Guinea, Nation. Sci. Mus.(Tokyo) 102 (1975).1975. 18
Polystichum decoratum Maxon [7] Polystichum decoratum Maxon, Contr. U.S.Natl. Herb. 13:30, t. 3.1909. 23W
Polystichum deflexum Ching ex W.M.Chu[25] [166] Polystichum deflexum Ching ex W.M.Chu, Acta Bot. Yunnan. Supp. 5:47. fig. 27.1992. 02
Polystichum diaphanum (Zoll. & Moritz) Moore [0] Aspidium diaphanum Zoll. & Moritz, Nat. en Geneesk. Arch. 2:203.1845 18
Polystichum dielsii Christ [1] [166] Polystichum dielsii H. Christ, Bulletin de l'Académie internationale de géographie botanique 1906:238.1906. 02
Polystichum diffundens H.S.Kung & L.B.Zhang [25] Polystichum diffundens H.S.Kung & L.B.Zhang, Acta Bot. Yunnan. 17(1):27. figs. 1:4-6.1995. 02
Polystichum discretum (D. Don) J.Sm. [3] [9] [12] [13] [57] [294] Aspidium discretum D. Don, Prod. Fl. Nepal. 41825. 02 16
Polystichum disjunctum Ching ex W.M. Chu & Z.R. He Polystichum disjunctum Ching ex W.M. Chu & Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):18. Pl. 3. f.6-9.2001 02
Polystichum dissimulans Maxon [56] Polystichum dissimulans Maxon, Contr. U.S.Natl. Herb.13:3l t.4A.1909 23W
Polystichum distans Fournier Polystichum distans Fournier, Mex.Pl. 1:911872. 23M
Polystichum dracomontanum Schelpe & N.C. Anthony [27] Polystichum dracomontanum Schelpe & N.C. Anthony, Contr. Bolus Herb. 10:145.1982. 11
Polystichum drepanum[0] Aspidium drepanum Sw., Schrad. Journ.1800(2):38.1801 05
Polystichum dubium (H.Karst.) Diels Phegopteris dubia H.Karst., Fl. Col. 1:169. t 84.1861. 23M, 27
Polystichum dudleyi Maxon [3] Polystichum dudleyi Maxon, Journ. Wash. Acad. Sci. 8:620.1918. 09
Polystichum duthiei (Hope) C.Chr.[0] [117] [166] [295] Aspidium duthiei Hope, Journal of the Bombay Natural History Society 12: 532 t. 61899. 02 16
Polystichum echinatum (Gmel.) C.Chr. [0][56] Polypodium echinatum J.F.Gmel. in L., Syst.Nat. 2(2): 1309.1791. 23W
Polystichum edanioi Copel. [4] Polystichum edanioi Copel., Philip. Journ. Sci. 83:97. t.2.1954. 18
Polystichum ekmani Maxon [56] Polystichum ekmanii Maxon, Proc. Biol. Soc. Wash. 46: 108.1933. 23W
Polystichum elevatovenosum Ching ex W.M. Chu and Z.R. He [166] Polystichum elevatovenosum Ching ex W.M. Chu and Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):148. Pl. 43. f. 6-7.2001. 02
Polystichum elmeri Copel.[4] Polystichum elmeri Copel., Philip. Journ. Sc. 81:21.1952. 18
Polystichum eriorachis v.A.v.R. [3] Polystichum eriorachis v.A.v.R., Bull. Buit. III.5:220.1922. 18
Polystichum erosum Ching & Shing[5] Polystichum erosum Ching & K.H.Shing, Acta Phytotax. Sin. 10:303.1965. 02
Polystichum erythrosorum A.R. Smith [db] Polystichum erythrosorum A.R.Smith, Proc.Calif.Acad.Sci. Ser.IV.40: 225.1975. 23M
Polystichum exauriforme H.S.Kung & L.B.Zhang [25] Polystichum exauriforme H.S.Kung & L.B.Zhang, Acta Bot. Yunnan. 17(1):27. fig. 1:7.1995. 02
Polystichum excelcior Ching & Z.Y.Liu [27] Polystichum excelcior Ching & Z.Y.Liu, Bull.Bot.Res., Harbin 4(4):16. f. 44.1984. 02
Polystichum excellens Ching [4] Polystichum excellens Ching, Icon. Fil. Sin. 5: t.234.1958. 02
Polystichum eximium(Mett. ex Kuhn)C.Chr. [3] [47] [117] Phegopteris eximium Mett. ex Kuhn, Linnaea 36:107.1869. 02 16 17
Polystichum falcatilobum Ching ex W.M. Chu & Z.R. He [166] Polystichum falcatilobum Ching ex W.M. Chu & Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):152. Pl. 44. f. 3-4. 02
Polystichum falcinellum(Sw.) C. Presl [0] Aspidium falcinellum Sw., Syn. 46,243.1806. 05
Polystichum fallax Tinsdale[4] Polystichum fallax Tindale, Proc. Linn. Soc. N.S. Wales 80:54. t.2.1955. 29
Polystichum fibrillopaleaceum(Kodama)Tagawa [21] Polystichum polyblepharum(Roem.)Presl var. fibrillopaleaceum (Kodama)Tagawa [47] Polystichum aculeatum (L.)Roth var. fibrillopaleaceum Kodama, Bot. Mag. Tokyo 29:??1915 02 JPN ONLY
Polystichum filiorum Mickel in Johns [56] Polystichum filiorum Mickel in Johns, Johns, R.J. (Ed.) Holttum Memorial Volume 127.1997. 23W
Polystichum fimbriatum H. Christ [1] [166] Polystichum fimbriatum H. Christ, Bulletin de l'Académie internationale de géographie botanique 16(205-206):237.1906. 02
Polystichum flaccidum v.A.v.R. [3] Polystichum flaccidum v.A.v.R., Bull. Buit. III. 5:220.1922. 18
Polystichum foenisecii (Lowe)P. Fourn. [4] Nephrodium foenisecii Lowe, Tr. Cambr. Phil. Soc. 4:7.1830. ??
Polystichum formosanum Ros. [2]; [233] Polystichum formosanum Ros., Hedwigia 56:338.1915. 02 TWN ONLY
Polystichum formosum Tindale[4] Polystichum formosum Tindale, S. Austr. Nat. 28:32.1954. 29
Polystichum fournieri A.R. Smith Polystichum fournieri A.R.Smith, Amer. Fern J. 70(1): 27.1980. 23M
Polystichum frigidicola H.S.Kung & L.B.Zhang [25] [166] Polystichum frigidicola H.S.Kung & L.B.Zhang, Acta Bot. Yunnan. 17(1):28. figs. 2:1-2.1995. 02
Polystichum fuentesii Espinosa [4:254] [239] Polystichum fuentesi Espinosa, Rev. Chil.Hist.Nat. 36: 239.1932. 20
Polystichum fugongense Ching & W.M.Chu ex W.M. Chu [166] Polystichum fugongense Ching & W.M.Chu ex W.M. Chu, Acta Bot. Yunnan. Supp. 5:49.1992. 02
Polystichum furfuraceum A.R. Smith [db] Polystichum furfuraceum A.R.Smith, Proc. Cal.Acad.Sci. Ser. IV. 40: 226,f6A-D1975. 23M
Polystichum fuscum Copel. [4] Polystichum fuscum Copel., Philip. Journ. Sc. 81:20. 1952. 18
Polystichum grandifrons C.Chr. [3] [57] Polystichum grandifrons C.Chr., Ind. Suppl. III:163. 1934. 02
Polystichum guadalupense Fée[0] [56] Polystichum guadalupense Fée, Mém. foug. 11:74 t.19 f.2. 1866. 23W
Polystichum guangxiense Chu and Zhou [166] Polystichum guangxiense W.M. Chu and H. G. Zhou, Guihaia 16(3): 205. f. 2. 1996. 02
Polystichum gymnocarpium Ching ex W.M. Chu and Z.R. He [166] Polystichum gymnocarpium Ching ex W.M. Chu and Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):146. Pl. 35. f. 5-6.2001. 02
Polystichum habaense Ching & H.S. Kung [27] Polystichum habaense Ching & H.S. Kung, Bull. Bot. Res. Harbin 9(3):41989. 02
Polystichum habbemense Nakaike [27] Polystichum habbemense Nakaike, Bull. Nat. Sci. Mus., Tokyo, B 2(2):51.1976. 18
Polystichum haleakalense Brack. [3,db] Polystichum haleakalense Brack., Expl. Exp. 16:204 t.28.1854. 21
Polystichum hancockii(Hance)Diels [0] [9] [21] [117] [166] Ptilopteris hancockii Hance, JoB. 1884:139.1884. 02
Polystichum harpophyllum (Zenker ex Kunze)Sledge [27] [57] Polypodium harpophyllum Zenker ex Kunze, 16
Polystichum harrisii Maxon Polystichum harrisii Maxon , Contr.U.S.Natl.Herb.13: 32 t.4B.1909 23W
Polystichum hartwegii(Klotzsch)Hieron. [db] Aspidium hartwegii Klotzsch, Linnaea 20: 366.1847 23M
Polystichum hecatopterum Diels [0] [21] [34] [12] [117] [166] [295] Polystichum hecatopterum Diels, Engl. Jahrb. 29:193.1900. 02
Polystichum herbaceum Ching & Z.Y.Liu[27] [166] Polystichum herbaceum Ching & Z.Y.Liu, Bull.Bot.Res., Harbin 4(4):20. f. 49.1984. 02
Polystichum heterolepis Fée [56] Polystichum heterolepis Fée, Gen. Fil. 2791850-52 23W
Polystichum hillebrandii Carr. in Seem. [3] Polystichum hillebrandii Carruthers in Seem., Fl. Vit. 358. 1873. 21
Polystichum holttumii C.Chr. in Holttum[3] [214] Polystichum holttumii C.Chr., Gardens Bull. S.S. 7:256 t. 53. 1934. 18
Polystichum hottense C.Chr. [4] [56] Polystichum hottense C.Chr., Kongl.Svenska Vetenskapsakad. Handl. ser.3.16(2):41 .t.12(1-3).1937. 23W
Polystichum houchangense Ching ex P.S.Wang [1666] Polystichum houchangense Ching ex P.S. Wang, Guizhou Sci. 12(2): 55. f. 1. 1994. 02
Polystichum huae H.S.Kung & L.B.Zhang [25] [166] Polystichum huae H.S.Kung & L.B.Zhang in Zhang & Kung, Acta Phytotax. Sin. 33(3):311. f. 2, C-G.1995. 02
Polystichum ichangense [0] [166] [295] Polystichum ichangense H. Christ, Bull. Soc. Fr. 52 Mém. I.28.1905. 02
Polystichum igaense Tagawa [4] [21] [47] Polystichum igaense Tagawa, Acta Phytotax. Geobot. 15:15.1953. 02 JPN ONLY
Polystichum ilicifolium Fée [3] [56] Polystichum ilicifolium Fée, Gen. Fil. 279.= Mém. foug. 5: 279.1852. 23W
Polystichum imbricans(D.C.Eaton)D.H.Wagner [27] Aspidium munitum Kaulf. var. imbricans D.C.Eaton, 04
Polystichum inadae Kurata [5] Polystichum inadae Kurata, J. Geobot. 11:6.1962. 02 JPN ONLY
Polystichum incisopinnulum H.S.Kung & L.B.Zhang [25] [166] [295] Polystichum incisopinnulum H.S.Kung & L.B.Zhang, Acta Bot. Yunnan. 17(1):25. fig. 2:31995. 02
Polystichum incongruum J.P. Roux Polystichum incongruum J.P. Roux, Bot. J. Linn. Soc. 125:36,37.1997. 11 (28?)
Polystichum integrilimbum Ching & H.S.Kung [27] [166] [295] Polystichum integrilimbum Ching & H.S.Kung, Acta Bot. Boreal-Occid. Sin. 9(4):275.1989. 02
Polystichum integrilobum (Ching ex Y.T. Hsieh) W.M. Chu ex H.S. Kung [166] [295] Sorolepidium integrilobum Ching ex Y.T. Hsieh, Bull. Bot. Res., Harbin 9(3): 48.1989. 02
Polystichum jinfoshanense Ching & Z.Y.Liu[27] Polystichum jinfoshanense Ching & Z.Y.Liu, Bull. Bot. Res. Harbin 3(4):29.1983. 02
Polystichum jiulaodongense W.M. Chu & Z.R. He [166] Polystichum jiulaodongense W.M. Chu & Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):227. 02
Polystichum kadrovii A. Askerov & A. Bobrov [5] Polystichum kadryovii A. Askerov & A. Bobrov, Bot. Zhurn. 57:1297.1972. 01 or 08 CAUC
Polystichum kalambatitrense Tardieu [4] Polystichum kalambatitrense Tardieu, Notul. Syst. [Paris] 15:165. f.1(1-4).1956. 15
Polystichum kangdongense H.S. Kung & L.B. Zhang [166] Polystichum kangdongense H.S. Kung & L.B. Zhang, Acta Phytotax. Sinica 36(3): 254. f. 3: 1-4.1998. 02
Polystichum keysserianum Ros. [1] Polystichum keysserianum Rosenst., Fedde Repert. 10:331.1912. 18
Polystichum kilimanjaricum Pichi-Serm. [5] Polystichum kilimanjaricum Pichi-Serm., Webbia 27:445. 1972. 12
Polystichum killipii Maxon [3]; P. X killipii [56] Polystichum killipii Maxon, Cont.U.S.Natl.Herb.24:53 t. 20.1922. 23W
Polystichum kinabaluense C.Chr. in Holttum [3] [214] Polystichum kinabaluense C.Chr. in Holttum, Gardens Bull. S.S. 7:255 t.52.1934. 18
Polystichum kruckebergii W.Wagner [5] Polystichum kruckebergii W. Wagner, Amer. Fern J. 56: 44, t,1.1966. 04
Polystichum krugii Maxon Polystichum krugii Maxon, Proc.Biol.Soc.Wash.18:215.1905 23W
Polystichum kurokawae Tagawa [4] [47] Polystichum kurokawae Tagawa, Journ. Jap. Bot. 26:20.1951. 02 JPN ONLY
Polystichum kwakiutlii D.H. Wagner [27] Polystichum kwakiutlii D.H.Wagner, Amer. Fern J. 80(2):50.1990. 04
Polystichum kwangtungense Ching [4] [166] [295] Polystichum kwangtungense Ching, Icon. Fil. Sin. 5: t.236.1958. 02
Polystichum lachenense(Hk.)Bedd. [57] Polystichum aleuticum C.Chr. ex Hultén, Svensk. Bot. Tidskr. 30:515. f.1(a).1936. 04
Polystichum lachenense(Hk.)Bedd.[0] [47] [57] [117] [166] Aspidium lachenense W.J.Hooker, Hk. sp. 4:8 t. 212.1862. 02 16
Polystichum lanceolatum (Baker) Diels [0] [166] Aspidium lanceolatum Baker, Gard. Chr. n.s. 14:494.1880. 02
Polystichum lancilobum C.Chr. [56] Polystichum lancilobum C.Chr., Kungl. Svenska Vetenspsakad. Handl.ser.3.16(2):40.t.9(1-4).1937. 23W
Polystichum langchungense Ching ex H.S. Kung [166] Polystichum langchungense Ching ex H.S. Kung, Chin. J. Appl. Environ. Biol. 3(2): 134. f. I, 1. 1997. 02
Polystichum latilepis Ching & H.S. Kung [27] [166] Polystichum latilepis Ching & H.S. Kung, Acta Boreal-Occid. Sin. 9(4):273. f.2: 3-4.1989. 02
Polystichum lehmannii Hier. Polystichum lehmannii Hieron., Engl. Jahrb. 34:452.1904. 23M, 27
Polystichum lemmonii Und.[0] Polystichum lemmoni Underw., Our Nat. Ferns ed. VI. 116.1900. 04
Polystichum lentum (D. Don) T. Moore [0] [57] [166] Aspidium lentum D. Don, Prod. Fl. Nepal. 4.1825. 16
Polystichum lepidotum M.A. Sundue & M. Kessler [321] Polystichum lepidotum M.A. Sundue & M. Kessler, Brittonia 57(3): 210. f. 1.2005. 27
Polystichum leveillei C.Chr.[2] [166] Polystichum leveillei C.Chr., Bulletin de l'Académie internationale de géographie botanique 1913:143.1913. 02
Polystichum lilianiae Barrington [322] Polystichum lilianae Barrington, Brittonia 55(4): 321. f. 2. 2003. 23M
Polystichum lindseaefolium Scort. in Ridley [3] Polystichum lindseaeafolium Scort. in Ridley, Journ. Mal. Br. R. As. Soc. 4:61. 1926. 17, 18
Polystichum lineare(C.Chr.)Copel. [4] [212] Papuapteris linearis C.Chr., Brittonia 2:300 f.2.1937. 18
Polystichum liuii Ching [27] [166] [295] Polystichum liuii Ching in Ching & Liu, Bull. Bot. Res. Harbin 3(4):28. 1983. 02
Polystichum lonchitis(L.)Roth [0] [47] Polypodium lonchitis L., sp. 2:1088.1753. 02 etc
Polystichum longifrons Kurata [5] Polystichum longifrons Kurata, Sci. Rep. Yokosuka City Mus. 10:31.1964. 02 JPN ONLY
Polystichum longipaleatum H. Christ [1] [57] [166] [295] Polystichum longipaleatum H. Christ in Lecomte, Not. syst. 1:35.1909. 02
Polystichum longipes Maxon [db] Polystichum longipes Maxon, Contr.U.S.Natl Herb. 13.34,t.6.1909. 23W
Polystichum longipinnulum Nair [5] Polystichum longipinnulum Nair, Amer. Fern J. 64:15. f. 1.1974. 16
Polystichum longispinosum Ching ex L.B. Xhang and H.S. Kung[166] Polystichum longispinosum Ching ex L.B. Xhang and H.S. Kung, Acta Phytotax. Sinica 36(5): 467. f. 1. 1998. 02
Polystichum longissimum Ching & Z.Y.Liu [27] [166] [295] Polystichum longissimum Ching & Z.Y.Liu, Bull. Bot. Res. Harbin 4(4)16. f. 45. 1984. 02
Polystichum luctuosum(Kze.)Moore[0 raised to sp.] [3] [57] Aspidium luctuosum Kunze , Linnaea 10:548.1836. 11 15
Polystichum macleae [13] Aspidium macleaii Baker in Hk., Ic. pl. t. 1654.1886. 11
Polystichum maevaranense Tardieu [4] Polystichum maevaranense Tardieu, Notul. Syst. [Paris] 15:166. f.1(5-9). 1956. 15
Polystichum magnificum Ballard [4] [221] Polystichum magnificum Ballard, Kew Bull. 1957:48. f.1.1957. 12
Polystichum makinoi (Tagawa)Tagawa[4:255] [12] [47] [57] [166] [295] Polystichum aculeatum (L.)Roth var. makinoi Tagawa, Acta Phytotax. Geobot. 1:88. 1932. 02
Polystichum manmeiense (H.Christ) Nakaike [57] [166] [295] Aspidium manmeiense H.Christ, Bull. Boiss. 6:965.1898 02
Polystichum mannii Hope ex Fraser-Jenkins [57] Polystichum mannii Hope ex Fraser-Jenkins, Asp. Pl. Sci. 13: 263.1991. 02
Polystichum marquesense E. Brown [3] Polystichum marquesense E. Brown, Bishop Mus. Bull. 89:40 t. 61931. 20
Polystichum martinii H. Christ [0] [57] [166] [295] Polystichum martini H. Christ, Bulletin de l'Académie internationale de géographie botanique 1902:263 c. tab.1902. 02
Polystichum mashikoi Kurata [4] Polystichum mashikoi Kurata, Journ. Geobot. 7:116.1958. 02 JPN ONLY
Polystichum maximum M. Kessler& A.R.Sm. [328] Polystichum giganteum M. Kessler & A.R.Sm., Brittonia 57(3): 215. f. 2C.2005. 27
Polystichum mehrae Fraser-Jenk. & Khullar [27] [57] [166] [295] Polystichum mehrae Fraser-Jenk. & Khullar, Indian Fern J. 2(1-2): 101985. 02 16
Polystichum meiguense Ching & H.S. Kung [27] [166] [295] Polystichum meiguense Ching & H.S. Kung, Acta Bot. Boreal-Occid. Sin. 9(4):269.1989. 02
Polystichum melanostipes Ching & H.S. Kung [27] [166] [295] Polystichum melanostipes Ching & H.S. Kung, Bull. Bot. Res. Harbin 9(3):10. f. 3.1989. 02
Polystichum mexiae Copel. Polystichum mexiae Copeland, Univ. Calif. Publ. Bot. 19: 299. t. 55.1941. 27
Polystichum mickelii A.R.Smith Polystichum mickelii A.R.Smith, Amer. Fern J. 70(1): 25.1980 23M
Polystichum microchlamys(Christ)Kodama in Matsum.[3] [47] Aspidium microchlamys H. Christ, Bull. Boiss. 7:830.1899. 02
Polystichum microsorium Fée Polystichum microsorium Fée, Crypt. Vasc. Br. 1:125,t.40,f.1.1869 26
Polystichum mohrioides(Bory)C. Presl Aspidium mohrioides Bory, Mém.Soc.Linn.Paris 4: 597; Dup.Voy.Bot. 1: 267,t.35,f.1.1828. 33 34
Polystichum moluccense (Blume) T. Moore [57] Aspidium moluccense Blume, Enum. 168.1828 18
Polystichum montevidense(Spreng.)Rosenst. Polypodium montevidense Spreng., Linnaeus, Syst. Veget. ed. 16. 4(1): 59.1827. 26 27
Polystichum monticola N.C.Anthony & Schelpe[27] Polystichum monticola N.C.Anthony & Schelpe, Bothalia 15(3-4):554.1985. 11
Polystichum moorei H. [0] Polystichum moorei H. Christ, Arkiv för Bot. 4(12):3.1905. 29
Polystichum morii Hayata [3] [21] [34] [117] [166] [295] Polystichum morii Hayata, Ic. Pl. Formosa 7. 95 f. 58.1918. 02 TWN ONLY
Polystichum mucronifolium (Bl.) C. Presl [0 as sp.] [57] [295] Aspidium mucronifolium Blume, Enum. 164.1828 16 18
Polystichum multifidum(Mett.)Moore Aspidium multifidum Mett., Filices Lechlerianae 20,t.3, f.14,15,16,17.1856. 33
Polystichum münchii(Christ)C.Chr. Aspidium münchii H. Christ, Bull. Boiss. II.1: 1120.1901 23M
Polystichum munitum (Kaulf.) C. Presl [0] Aspidium munitum Kaulf., Enum. 236.1824. 04
Polystichum muricatum (L.) Liebm. Polypodium muricatum L., Species Plantarum 2. 1093.1753 23M, 23W, 27
Polystichum muscicola Ching ex W.M. Chu and Z.R. He Polystichum muscicola Ching ex W.M. Chu and Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):140. Pl. 45. f. 1-2.2001. 02
Polystichum muticum Copel. [4] Polystichum muticum Copel., Univ. Calif. Publ. Bot. 18:221.1942. 18
Polystichum myer-dreesii Copel. [4] Polystichum myer-dreesii Copel., Univ. Calif. Publ. Bot. 18:221.1942. 18
Polystichum nahuelhuapiense Diem [4] Polystichum nahuelhuapiense Diem, Darwiniana 12: 67, f. 1.1960. 33
Polystichum nayongense P.S. Wang & X.Y. Wang [166] [295] Polystichum nayongense P.S. Wang & X.Y. Wang, Acta Bot. Yunnan 19(1):41. f. 1. 1997. 02
Polystichum neolobatum Nakai [3] [21] [12] [47] [117] [166] Polystichum neo-lobatum Nakai, Bot. Mag. Tokyo 39:118.1925. 02
Polystichum neozelandicum Fée [249] [266] Polystichum neozelandicum Fée,
Polystichum neozelandicum ssp. zerophyllym (Colenso) L. Perrie Aspidium zerophyllum Col., New Zealand1897. 35
Polystichum nepalense (Spreng.) C.Chr. [0] [117] Aspidium nepalense Spreng. , Syst. 4: 97.1827. 02 16 18
Polystichum nigrum Ching & H.S. Kung [27] [166] [295] Polystichum nigrum Ching & H.S. Kung in Kung, Bull. Bot. Res. Harbin 9(3):8. f. 2.1989. 02
Polystichum ningshenense Ching & Hsu[27] [166] [295] Polystichum ningshenense Ching & Hsu, Fl. Tsinlingensis 2:223. t. 39. f. 3-9. 1974. 02
Polystichum nudicaule Rosenst. Polystichum nudicaule Rosenst., Fedde Repert. 11: 56.1912. 27
Polystichum nudum Copeland [1] Polystichum nudum Cop., Phil. Journ. Sci. 1: Suppl. II. 145.1906. 18
Polystichum obai Tagawa [3] Polystichum obai Tagawa, Acta Phytotax. 2:194.1933. 02 JPN ONLY
Polystichum obliquum(D. Don)T. Moore [0] [21] [117] Aspidium obliquum D. Don, Prod. Fl. Nepal. 3.1825. 02 16 18
Polystichum oblongum Ching ex W.M. Chu and Z.R. He [166] Polystichum oblongum Ching ex W.M. Chu and Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):132, 228.2001 02
Polystichum obtusum J. Sm. ex C. Presl [0] [214] Aspidium obtusum J. Sm. ex C. Presl, Epim. 53.1849. 18
Polystichum oculatum (Hook.) J. B. Armstr. [249] [266] Aspidium oculatum W.J.Hooker, Species filicum 4:24 t. 228. 1862. 35
Polystichum ohmurae Kurata [4] Polystichum ohmurae Kurata, Hokuriku Journ. Bot. 5:79. 1956. 02 JPN ONLY
Polystichum oligocarpum Ching ex H.S.Kung & L.B.Zhang [25] [166] [295] Polystichum oligocarpum Ching ex H.S.Kung & L.B.Zhang, Acta Phytotax. Sin. 33(3):308. f. 1, A-D.1995. 02
Polystichum omeiense C.Chr.[0] [12] [166] Polystichum omeiense C.Chr., Ind. Fil. 67. 1905; 585. 1906.1905. 02
Polystichum orbiculatum (Desv.) Remy & Fée in Gay [db] Aspidium orbiculatum Desv., Berl.Mag. 5:321.1811. 23M 27 PAL
Polystichum ordinatum(Kze.)Liebm. [db] Aspidium ordinatum Kunze , Linnaea 18:347.1844. 23M
Polystichum oreodoxa H.S.Kung & L.B.Zhang ex Ching [25] [166] [295] Polystichum oreodoxa Ching ex H.S.Kung & L.B.Zhang, Acta Phytotax. Sin. 33(3):309. f. 2 A,B.1995. 02
Polystichum otomasui Kurata [5] Polystichum otomasui Kurata , J. Geobot. 12:67. Hokuriku J. Bot. 12:67.1963. 02 JPN ONLY
Polystichum otophorum(Franch.)Bedd.[0] [166] Aspidium otophorum Franch., Nouv. Arch. Mus. II.10:116. 1888. 02
Polystichum ovatopaleaceum(Kodama)Kurata [5] Polystichum aculeatum (L.)Roth var. ovatopaleaceum Kodama, Bot. Mag. Tokyo 29:323.1915. 02
Polystichum paleatum Copel. [4] [db] Polystichum paleatum Copel., Occ. Pap. Bishop Mus. 14:58. t.9. 1938. 20
Polystichum papuanum C.Chr. [4] Polystichum papuanum C.Chr., Brittonia 2:299. 1937. 18
Polystichum paramicola Kessler& A.R.Sm. [321] Polystichum paramicola M. Kessler & A.R.Sm., Brittonia 57(3): 222. f. 5C.2005. 27
Polystichum parvifoliolatum W.M.Chu [25] [166] [295] Polystichum parvifoliolatum W.M.Chu , Acta Bot. Yunnan. Supp. 5:49. f. 29.1992. 02
Polystichum parvipinnulum Tagawa [4] [117] [166] [295] Polystichum parvipinnulum Tagawa, Acta Phytotax. Geobot. 6:160.1937. 02 TWN ONLY
Polystichum pauciaculeatum Bonap. [3] Polystichum pauciaculeatum Bonap., N. Pt. 7:206.1918. 15
Polystichum peishanii Zhang & He Polystichum peishanii Zhang & He, Botanical Studies 50:102. f. 1,2. 2009. 02
Polystichum pianmaense W.M.Chu [25] [166] [295] Polystichum pianmaense W.M.Chu , Acta Bot. Yunnan. Supp. 5:51. f. 30.1992. 02
Polystichum piceopaleaceum Tagawa [4] [21] [57] [117] [166] [295] Polystichum piceo-paleaceum Tagawa, Acta Phytotax. Geobot. 5:255.1936. 02
Polystichum pilosum Copel. [4] Polystichum pilosum Copel., Journ. Arnold Arbor. 30:439.1949. 20
Polystichum plaschnickianum(Kze.)Moore [0] [56] Aspidium plaschnickianum Kunze , Linnaea 23: 229 [or 239, fide 56],3021850. 23W
Polystichum platylepis Fée [0 as sp.] Polystichum platylepis Fée, Cr.Vasc.Br. 1: 129,t.41[40 sic in text],f.11869 26
Polystichum platyphyllum(Willd.) C. Presl Aspidium platyphyllum Willd., Sp.Pl.ed.5. 5: 255.1810. 23M 23W 27 26
Polystichum polyblepharum(Röm. ex Kunze) C. Presl [47] [166] [295] Aspidium polyblepharon Römer ex Kunze, Bot. Zeit. 1848:572.1848. 02
Polystichum polyodon Wall. ex Ching [4] [57] Polystichum polyodon Wall. ex Ching, Bull. Chin. Bot. Soc. 2:93.1936. 16
Polystichum prescottianum (Wall. ex Mett.) T. Moore [0] [57] [117] [166] [295] Aspidium prescottianum Wall. nomen nudum, List n. 363.1828. 02 16
Polystichum prionolepis Hayata [2] [117] [166] [295] Polystichum prionolepis Hayata, Ic. Fl. Formosa 4:197 f. 134.1914. 02 TWN ONLY
Polystichum proliferum(R.Br.)Presl [0 as sp.] Aspidium proliferum R. Br., Prod. Fl. N. Holl. 1471810 29
Polystichum pseudoacutidens Ching ex W.M. Chu & Z.R. He [166] Polystichum pseudoacutidens Ching ex W.M. Chu & Z.R. He, Acta Botanica Yunnanica 22(3):257. t. 2. f. 3,4.2000 02
Polystichum pseudomakinoi Tagawa [4] [21] [47] [166] [295] Polystichum pseudo-makinoi Tagawa, Acta Phytotax. Geobot. 5:257. 1936. 02
Polystichum pseudomicrophyllum Nakaike [27] Polystichum pseudomicrophyllum Nakaike, Bull. Nat. Sci. Mus. Tokyo B 9(3): 103.1983. 18
Polystichum pseudorhomboideum H.S. Kung & L.B. Zhang [166] [295] Polystichum pseudorhomboideum H.S. Kung & L.B. Zhang, Acta Phytotax. Sinica 36(5): 468.1998. 02
Polystichum pseudosetosum Ching & Z.Y.Liu[27] [166] [295] Polystichum pseudo-setosum Ching & Z.Y.Liu, Bull.Bot.Res., Harbin 4(4):19. f. 48.1984. 02
Polystichum pseudoxiphophyllum Ching ex H.S. Kung [166] Polystichum pseudo-xiphophyllum Ching ex H.S. Kung, Chin. J. Appl. Environ. Biol. 3(2): 135. f. I: 2-3. 1997. 02
Polystichum pullenii Nakaike[27] Polystichum pullenii Nakaike, Bull. Nat. Sci. Mus. Tokyo B 43(3):129.1978. 18
Polystichum punctiferum C.Chr.[3] [57] [166] [215] Polystichum punctiferum C.Chr., Contr. U.S. Nat. Herb. 26:288 t. 171931. 02
Polystichum pungens(Klf.)Presl [0] Aspidium pungens Klf., Enum. 242.1824. 11
Polystichum pycnolepis(Kze. ex Kl.)Moore [rmt, db] Aspidium pycnolepis Kunze ex Klotzsch, Linnaea 20: 365.1847 27 PAL
Polystichum pycnopterum(H.Christ)Ching ex W.M.Chu & Z.R. He [166] Aspidium aculeatum var. pycnopterum H. Christ, Bull. Herb. Boiss. 7:14.1899. 02
Polystichum rachichlaena Fée [db] Polystichum rachichlaena Fée, Mém. Foug. 8: 100.1857 23M
Polystichum rapense E. Brown [3] Polystichum rapense E. Brown, Bishop Museum Bull. 89: 39 t. 4A.1931. 20
Polystichum retrosopaleaceum(Kodama)Tagawa [47] [166] [295] Polystichum aculeatum var. retroso-paleaceum Kodama, Bot. Mag. Tokyo 29: (330)1915. 02
Polystichum revolutum P.S. Wang in Wang & Wang Polystichum revolutum P.S. Wang in Wang & Wang, Pteridophyte Flora of Guizhou 549. t. 133. f. 6-12.2001. 02
Polystichum rhizophyllum(Sw.)Presl [0] Polypodium rhizophyllum Sw., Prodr. veg. ind. occ. 132.1788 23W
Polystichum rigens Tagawa [4] [21] [47] [166] [295] Polystichum rigens Tagawa, Acta Phytotax. Geobot. 6:91.1937. 02
Polystichum robustum Ching ex L. B. Zhang & H. S. Kung Polystichum robustum Ching ex L. B. Zhang & H. S. Kung, Acta Phytotax. Sinica 36(5): 468. f. 2-5. 1998. 02
Polystichum rochaleanum Fée Polystichum rochaleanum Fée, Cr. Vasc. Br. 2: 69, t. 99, f. 2.1872-73 26
Polystichum rufopaleaceum Ching ex H. S. Kung and L. B. Zhang [166] [295] Polystichum rufopaleaceum Ching ex H. S. Kung and L. B. Zhang, Acta Phytotax. Sinica 36 (3): 246. f. 1. 1998. 02
Polystichum rufum M. Kessler & A.R.Sm. [321] Polystichum rufum M. Kessler & A.R.Sm., Brittonia 57(3): 217. f. 2D.2005. 27
Polystichum rupicola W.M.Chu ex Ching [25] [166] [295] Polystichum rupicola Ching ex W.M.Chu , Acta Bot. Yunnan. Supp. 5:51. f. 31.1992. 02
Polystichum salwinense Ching & H.S. Kung [166] [[295] Polystichum salwinense Ching & H.S.Kung, Bull. Bot. Res., Harbin 9(3):7. f. 5.1989. 02
Polystichum saxicola H.S. Kung & L.B. Zhang [166] [295] Polystichum saxicola H.S. Kung & L.B. Zhang, Acta Phytotax. Sinica 36(3): 250. f. 2.1-4. 1998. 02
Polystichum scariosum(Roxb.)C. Morton [5] [41] [57] Polypodium scariosum Roxb. , Calc. Journ. 4:494.1844. 18 02
Polystichum scopulinum (D.C.Eaton) Maxon Aspidium aculeatum var. scopulinum D.C. Eaton, Ferns N. Am. 2. 125 t. 62 f. 81880. 01 04
Polystichum semifertile(Clarke)Ching [4] [57] [166] [295] Aspidium aculeatum (L.) Sw. var. semifertile Clarke, Trans. Linn. Soc. London. ser. 2. Bot. 1:509.1880. 16, 17
Polystichum setiferum (Forssk.) T. Moore ex Woynar Polypodium setiferum Forssk., Flora Aegyptiaco-Arabica 185. 1775. 06 05
Polystichum setigerum Presl [db] Polystichum setigerum Presl, Tent. 83.1836. 04
Polystichum setillosum Ching [3] [166] Polystichum setillosum Ching, Bull. Fan Mem. Inst. 2:188. t. 3.1931. 02
Polystichum shandongense J.X.Li & Y. Wei [27] [166] [295] Polystichum shandongense J.X.Li & Y. Wei, Acta Phytotax. Sin. 22(2):164. f. 1.1984. 02
Polystichum shensiense H. Christ [1] [57] [166] [295] Polystichum shensiense H. Christ, Bulletin de l'Académie internationale de géographie botanique 1906:113.1906. 02
Polystichum shimurae Sa.Kurata ex Seriz. [27] Polystichum shimurae Sa.Kurata ex Seriz., J.Jap.Bot. 54(5):139. f.1. 1979. 02
Polystichum silaense Ching [4] [295] Polystichum silaense Ching, Bull. Fan. Mem. Inst. Biol. Bot. 11:71.1941. 02
Polystichum silvaticum(Colenso)Diels [176] [218] Polypodium silvaticum Colenso, Tasm. Journ. 2:163.1844. 35
Polystichum simplicius(Makino)Tagawa [3] Aspidium aristatum var. simplicius Makino, Bot. Mag. Tokyo 15:61.1901. 02 JPN ONLY
Polystichum sinense H. Christ [166] [295] Polystichum sinense H. Christ var. sinense , Bull. Soc. Fr. 52 Mém. 1:30. 1905. 02
Polystichum sodiroi H. Christ Polystichum sodiroi H. Christ, Arkiv för Bot. 4(12):4.1905 27
Polystichum solomonii Kessler& A.R.Sm. [321] Polystichum solomonii M. Kessler & A.R.Sm., Brittonia 57(3): 220. f. 5B.2005. 27
Polystichum sozanense H.S.Kung & L.B.Zhang ex Ching [25] [166] [295] Polystichum sozanense Ching ex L.B.Zhang & H.S.Kung, Acta Phytotax. Sin. 33(3):309. f. 1, E-I.1995. 02 TWN ONLY
Polystichum speciosissimum(A. Braun ex Kunze)R.M.Tryon & A.F. Tryon [db] Cheilanthes speciosissima A. Braun ex Kunze, Anal. Pterid. 35. t. 23. 1837. 23M
Polystichum spongiosum Maxon Polystichum spongiosum Maxon, Amer. Fern J. 19: 47.1929. 23W
Polystichum squarrosum(D.Don)Fée [0 as sp.] [3] [57] [166] [295] Aspidium squarrosum D. Don, Prod. Fl. Nepal. 4. 1825. 16 02
Polystichum stimulans [3] [db] [57] [166] Polystichum stimulans (Kunze ex Mett.) Bedd., Ferns Brit. India t. 31.1865. 02 16
Polystichum stokesii E.Brown [3] Polystichum stokesii E. Brown, Bishop Mus. Bull. 89:40. t.5.1931. 20
Polystichum stübelii Hieron. Polystichum stuebelii Hieron., Hedwigia 46:355,t8,f21,21a,21b1907 27
Polystichum subacutidens L.L.Xiang ex Ching [25] [166] [295] Polystichum subacutidens Ching ex L.L.Xiang, Acta Phytotax. Sin. 32(3):261. f. 2.1-3.1994. 02
Polystichum subapiciflorum Hayata [2] [57] Polystichum subapiciflorum Hayata, Ic. Fl. Formosa 5:335 f. 140. 1915. 02 16 17 18
Polystichum subdeltodon Ching [4] [166] Polystichum subdeltodon Ching, Bull. Fan Mem. Inst. Biol. ser.2. 1:308.1949. 02
Polystichum subfimbriatum W.M. Chu & Z. R. He [166] [295] Polystichum subfimbriatum W.M. Chu & Z. R. He, Acta Phytotax. Sinica 36 (4): 382. f. 1: 1-6. 1998.
Polystichum subinerme (Kze.) Fraser-Jenkins [57] Aspidium subinerme Kunze , Linnaea 24. 290.1851. 16
Polystichum subintegerrimum (Hook. & Arn.) R.Rodr. [[db - see notes on type]] Aspidium macrolepidum Desv., Prod. 248. 1827. 33
Polystichum subintegerrimum (Hook. & Arn.) R.Rodr. [db] Aspidium subintegerrimum Hook. & Arn., Bot.Beechey's Voyage 521832 33
Polystichum submarginale(Baker)Ching ex P.S. Wang [166] Polystichum auriculatum (L.) C. Presl var. submarginale Baker, Journ. Bot. 26:227.1888. 02
Polystichum subtripteron Tzvelev [27] Polystichum subtripteron Tzvelev, Novosti Sist. Vÿssh. Rast. 26:7.1989. 01?
Polystichum subulatum L.B.Zhang ex Ching [25][166] [295] Polystichum subulatum Ching ex L.B.Zhang , Acta Bot. Yunnan. 16(2):133. f. 1.2, 1.3.1994. 02
Polystichum tacticopterum (Kze.) Moore [166] [295] Aspidium tacticopterum Kunze , Linnaea 24:290.1851. 02 16
Polystichum tagawanum Kurata [4] [21] Polystichum tagawanum Kurata, Journ. Geobot. 7:41.1958. 02 JPN ONLY
Polystichum taizhongense H.S.Kung [166] [295] Polystichum taizhongense H. S. Kung , Acta Phytotax. Sinica 35(5): 461. f. 1. 1997. 02
Polystichum takakii Nakaike [5] Polystichum takakii Nakaike, Bot. Exped. Papua New Guinea, Nation. Sci. Mus. (Tokyo): 111.1975. 18
Polystichum talamancanum Barrington [db] Polystichum talamancanum Barrington, Ann. Missouri Bot. Gard. 76(2):369.1989. 23M
Polystichum tetragonum Fée Polystichum tetragonum Fée, 8 mem. 99. 1857.1857 33
Polystichum thomsonii (Hook.f. in Hook.)Bedd. [0] [21] [117] [166] [295] Aspidium thomsoni Hook.f. in Hook.), 2 Cent. t. 25.1860. 02 16
Polystichum tortuosum Nakaike [27] Polystichum tortuosum Nakaike, Bull. Nat. Sci. Mus. Tokyo B 2(3):102.1976. 18
Polystichum transkeiense W. Jacobsen [27] Polystichum transkeiense W.Jacobsen, J.S.African Bot. 44(2):169.1978. 28 11 12?
Polystichum transvaalense N.C.Anthony [27] Polystichum transvaalense N.C.Anthony, Contr. Bolus Herb. 10:146.1982. 11 12?
Polystichum trapezoides(Sw.)Presl [56] Aspidium trapezoides Sw., J.Bot. Schrad. 1800(2):31.1801. 23W
Polystichum triangulum(L.)Fée Polypodium triangulum L., Sp.Pl.2: 1088.1753 23W
Polystichum tridens(Hk.)Fée [0] Aspidium tridens Moore ex W.J.Hooker, Sp. Fil. 4: 15. t. 215.1862 23W
Polystichum tripteron(Kze.)Presl [0][12] [47] [166] [295] Aspidium tripteron Kunze, Bot. Zeit. 1848:569.1848. 02
Polystichum tsangmaiense H.S.Kung & Y.Tateishi [25] [166] [295] Polystichum tsangmaiense H.S.Kung & Y.Tateishi, Acta Phytotax. Sin. 32(3):268. Fig. 1.1-1.5.1994. 02 TIBET ONLY
Polystichum tsaratananense Tardieu[4] [246] Polystichum tsaratananense Tardieu, Mém. Inst.Sc. Madagascar ser. B. 7:44. 1956. 15
Polystichum tsingkanshanense Ching ex K.H.Shing & J.F.Cheng [27] Polystichum tsingkanshanense Ching ex K.H.Shing & J.F.Cheng, Jiangxi Sci. 8(3):46.1990. 02
Polystichum tsus-simense (Hook.) J.Sm. [0 as sp.] [3:165] [12] [47] [117] [166] Aspidium tsus-simense Hook., Species filicum 4:16 t.220.1862. 02
Polystichum turrialbae Christ Polystichum turrialbae H. Christ, Bull. Herb. Bois. II.6: 163.1906. 23M 27
Polystichum underwoodii Maxon [1] Polystichum underwoodii Maxon, Contr.U.S.Natl.Herb. 13: 38. t. 9.1909 23W
Polystichum vestitum (G. Forst.) C. Presl [0] [176] [218] Polypodium vestitum Georgio Forster, Flor.Insularum Australium Prodromus. 82. Gottingae. Dieterich.1786. 34, 35
Polystichum viviparum Fée Polystichum viviparum Fée, Gen. Fil. 280. 6 mém. 21 t. 3 f. 31850-52 23W
Polystichum volkensii Hieron. in Engl. [0] Aspidium volkensii Hieron. in Engl., Pflw. Ostafr. C. 86.1895. 12
Polystichum vulcanicum (Bl.) Moore Aspidium vulcanicum Blume, Enum. 163.1828. 18
Polystichum walkerae(Hook.)Sledge [228] Polypodium walkerae Hook. var. walkerae , Sp. Fil. 4:233.1862. 16
Polystichum wattii(Beddome)C.Chr. [0] [57] [166] [295] Aspidium wattii Bedd., JoB. 1888:34.1888. 02
Polystichum wawranum (Szyszyl. in Wawra) Perrie [249] [266] Aspidium wawranum Szyszyl., in Wawra, Itin. princ. Coburgi 126. t. 15 1888. 35
Polystichum weimingii Zhang and He Polystichum weimingii Zhang and He, Syst. Bot. 34(1): 13. f.1. 2009. 02
Polystichum whiteleggei Watts [2] Polystichum whiteleggei Watts, Proc. Linn. Soc. N.S. Wales 39 p. 257-262.1914. 29
Polystichum wilesianum Proctor [5] Polystichum wilesianum Proctor, Brit. Fern Gaz. 10:24, t. 3, 1968. 23W
Polystichum wilsonii H. Christ [221] Polystichum alticola Schelpe & N.C. Anthony, Contr. Bolus Herb. 10:144.1982. 11,12
Polystichum wilsonii H.Christ [1] [21] [13] [57] [117] Polystichum wilsonii H. Christ, Bot. Gaz. 51:353.1911. 02 10 11 15 16
Polystichum woronowii Fomin [1] [57] Polystichum woronowii Fomin, Moniteur Jard. bot. Tiflis livr. 18. 21. tab.1911. 01 or 08 CAUC
Polystichum wrightii(Baker)C.Chr. ex Maxon Polypodium wrightii Baker, Syn. Fil. 2: 304.1874. 23W
Polystichum xiphophyllum (Baker) Diels [0] [21] [117] [166] [295] Aspidium xiphophyllum Baker, JoB. 1888:227.1888. 02
Polystichum yadongense Ching & S.K.Wu [27] [166] [295] Polystichum yadongense Ching & S.K.Wu, Fl. Xizangica 1:220.1983. 02 TIBET ONLY
Polystichum yuanum Ching [4] [166] [295] Polystichum yuanum Ching, Icon. Fil. Sin. 5: t.235. 1958. 02
Polystichum yunnanense H. Christ [1] [57] [166] Polystichum yunnanense H. Christ, Not. syst. 1:34. 1909. 02 16
Polystichum zambesiacum Schelpe [5] Polystichum zambesiacum Schelpe, Bol. Soc. Brot. Sér. 2,41:215.1967. 12, 11
Polystichum zayuense Ching ex W.M. Chu & Z.R. He [166] Polystichum zayuense W.M. Chu & Z.R. He, Flora Reipublicae Popularis Sinicae vol. 5(2) Pteridophyta. Dryopteridaceae (2):157. Pl. 45. f.5-6.2001. 02

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