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Donovan, T. M. and J. Hines. 2007. Exercises in Occupancy Estimation and Modeling.

Chapter 16. Multiple-State Occupancy Models.

Another extension of the single-season, single-season occupancy model is the multiple-state model.In the Occupancy Estimation and Modeling book, the authors describe a scenario where individual ponds (sites) are monitored for evidence of breeding by amphibians, which might include finding egg masses, new metamorphs, or observing animals in amplexus. In this case, the site is classified into one of two states: it contains breeders or it contains non-breeders. The multi-state occupancy model is a straight-forward extension of the single-season occupancy model, and also incorporates some elements of the error model that we discussed in previous exercises.

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