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Effective conservation of forest songbirds requires knowledge of bird response
to landscape pattern, including information about reproductive success. Negative
impacts of forest fragmentation on avian productivity have been documented,
particularly in the highly fragmented agricultural landscapes of midwestern and mid-Atlantic
states. Relatively few studies have been conducted in extensively forested
landscapes. Rather than being fragmented, many forests in these landscapes are
characterized by a forest matrix perforated by scattered openings.

I conducted a landscape-level study of songbird productivity on the Green
Mountain National Forest (GMNF) in central Vermont. My
study area is an example of a
perforated, rather than fragmented, forest landscape. I compared songbird species
composition, abundance, and productivity on disturbed-canopy and closed-canopy
sites. The disturbed-canopy sites were characterized by canopy openings accounting
for 10% of site area, and were selected from a focal
analysis of the study area.

One paper from this study describes the use of line-transect surveys
to estimate density of fledgling broods. A second paper reports on the
investigation of altered landscape pattern as it relates to songbird
productivity.

Distance Sampling to Estimate Fledgling Brood Density of Forest Birds

Ernest W. Buford, David E. Capen, and B. K. Williams. 1996. Canadian
Field-Naturalist 110(4):642-648.
Abstract: Research on the status of avian communities often relies on estimates of abundance, but
does not always consider demographic factors such as productivity. We introduce the application
of a distance-sampling technique for estimating brood density of fledgling birds in forested
habitats. During 1993 and 1994, we conducted 60 line-transect surveys on 10 sites in the Green
Mountain National Forest in Vermont. Sites were divided into 2 groups, which allowed us to test
for differences in fledgling density between groups. We detected 508 broods representing 38
species. Using standard distance-sampling procedures, we estimated densities of 0.576 (C.V.
= 11.89) and 0.513 (C.V. = 12.54) broods per hectare. Density was not statistically different
between groups. As with other survey methods, distance sampling favors easily detected species;
however, line-transect density of fledglings is less obtrusive and less labor-intensive than
mist-netting or nest searches, and can contribute important information to studies of avian
communities.

Abundance and Productivity of Forest Songbirds in a Managed, Unfragmented Landscape in Vermont

Ernest W. Buford and David E. Capen. 1999. Journal of Wildlife Management 63(1):180-188.
Abstract: Effective conservation of
forest songbirds requires knowledge of responses to landscape pattern. We studied a breeding
songbird community in a landscape characterized by scattered openings in a forest matrix in Vermont.
We measured species composition, abundance, and fledgling brood density in the forested areas of
10 500-ha sites; we compared a group of 5 disturbed-canopy sites (<10% canopy
disturbance) with a control group of 5 undisturbed-canopy sites. We conducted 456 point counts
to determine species composition and abundance, and we conducted 60 line transect surveys to
estimate fledgling brood density as a measure of productivity. Adult densities were not different
between groups (P > 0.05) for 18 species listed as priority for conservation, although
power was low. Combined brood densities of all species, Neotropical migrants, and ground nesters
were not different among groups (P > 0.05). Brood densities of 3 neotropical migrants
also were not different (P > 0.05). However, combined brood density of area-sensitive,
forest interior species was greater on control sites (P < 0.05). At the scale of this
study, a minimal amount of canopy removal in the extensively forested landscape did not affect
abundance and overall productivity of songbirds inhabiting the remaining forest, but had begun
to suppress productivity of some forest interior species.
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